Related papers:
https://www.nature.com/articles/ng.3464
https://journals.plos.org/plosgenetics/article?id=10.1371/journal.pgen.1007908
https://www.biorxiv.org/content/10.1101/233486v2
https://onlinelibrary.wiley.com/doi/abs/10.1111/1755-0998.12747